Test free or more commonly attached, proloculus followed by slowly enlarging close-coiled tubular second chamber of one or two volutions, then becoming irregular in growth; wall agglutinated; aperture at the open end of the tube. U. Devonian (Frasnian); Aus­tralia; Germany. L. Carboniferous, L. Missis­sippian (Kinderhookian); USA: Indiana; to U. Carboniferous, U. Pennsylvanian; USA: Oklahoma, Indiana, Texas.